Absolutely solid entry. Good call on the palette swapping and really nice amount of content! The game compelled me to keep playing to see whats next all the way until the end. Some of the enemies felt a bit unfair in how they behaved or with their placement, but nothing too crazy to really hurt the gameplay. The attack timers seemed to function even when scrolling to a new screen giving the player no reaction time. I think the design decision to make certain boss attacks produce hearts was the right call, but at the same time it almost made some of the bosses feel too easy.
Is there some sort of extra ending for getting all the seeds. I think I collected all but maybe one.
This really felt like a complete package. Impressive work!