Lanak
Lanak | |
---|---|
Thesium scabrum | |
Sistematika | |
Carstvo: | Plantae |
Divizija: | Tracheophyta |
Razred: | Magnoliopsida |
Red: | Santalales |
Porodica: | Thesiaceae |
Rod: | Thesium L. |
Baze podataka | |
Lanak (lat. Thesium), biljni rod tradicionalno iz porodice santalovki[1]., međutim, Nickrent et al. (2010) smještaju Thesium u porodicu Thesiaceae. Nedavne taksonomske promjene proširile su rod uključivanjem prethodno opisanih rodova kao što su Kunkeliella Stearn i Thesidium Sond. (Forest i Manning 2013). Iako se Thesium može pronaći u Europi, Aziji, Australiji, Južnoj Americi i Sjevernoj Americi (ljudskim uvođenjem), velika većina vrsta su afričke, a najveća raznolikost nalazi se u regiji Cape. Doista, činjenica da su Lacomucinaea Nickrent & M.A.García i Osyridicarpos A.DC. sestrinski su Thesiumu (Nickrent i García 2015) i oba su ograničena na južnu Afriku, što ukazuje da je ovo vjerojatno područje odakle je rod nastao.[2]
Biom Fynbos, ugrađen u južnoafričku florističku regiju Cape, koja se ponekad naziva i florističko kraljevstvo (Takhtajan 1986), jedna je od floristički najraznolikijih regija na svijetu (Mucina i Rutherford 2006). Doista, od oko 170 imenovanih vrsta Thesium iz Južne Afrike, preko 40 od njih se nalazi samo u regiji Cape. Velika raznolikost komplicira identifikaciju svih kritosjemenjača, ali to je osobito istinito u slabo dokumentiranim rodovima kao što je Thesium. Posljednji veliki taksonomski radovi koji se bave Thesiumom u Južnoj Africi bili su od Hilla (1915., 1925.), ali od tada je imenovan niz novih vrsta. Ključevi predstavljeni u Hillovim radovima danas nisu osobito korisni u identificiranju južnoafričkog Thesiuma jer ne uključuju novije imenovane vrste, npr. one od Browna (1932.) ili Comptona (1931.). Štoviše, znakovi koji se koriste za razlikovanje vrsta nisu učinkoviti kada se ispituje širok spektar uzoraka. Ovo je pogoršano fenotipskom plastičnošću koja se vidi kod nekih vrsta u različitim razvojnim fazama i pod različitim ekološkim uvjetima (vatra, ispaša, nadmorska visina, supstrat itd.).[2]
Postoji preko 350 vrsta; po Kew-u 341. U Hrvatskoj također raste nekoliko vrsta (navedene su u popisu uz latinske nazive)[3].
- Thesium acuminatum A.W.Hill
- Thesium acutissimum A.DC.
- Thesium aellenianum Lawalrée
- Thesium affine Schltr.
- Thesium afghanicum Hendrych
- Thesium aggregatum A.W.Hill
- Thesium alatavicum Kar. & Kir.
- Thesium alatum Hilliard & B.L.Burtt
- Thesium albomontanum Compton
- Thesium alpinum L., planinski lanak
- Thesium amicorum Lawalrée
- Thesium angolense Pilg.
- Thesium angulosum A.DC.
- Thesium annulatum A.W.Hill
- Thesium annuum Lawalrée
- Thesium aphyllum Mart. ex A.DC.
- Thesium archeri Compton
- Thesium asperifolium A.W.Hill
- Thesium aspermontanum Zhigila, Verboom & Muasya
- Thesium asterias A.W.Hill
- Thesium atratum N.Lombard & M.M.le Roux
- Thesium atrum A.W.Hill
- Thesium aureum Jaub. & Spach
- Thesium auriculatum Vandas
- Thesium australe R.Br.
- Thesium bangweolense R.E.Fr.
- Thesium basninianum Turcz.
- Thesium bathyschistum Schltr.
- Thesium bavarum Schrank, gorski lanak
- Thesium bequaertii Robyns & Lawalrée
- Thesium bergeri Zucc.
- Thesium bertramii Azn.
- Thesium billardierei Boiss.
- Thesium boissieranum A.DC.
- Thesium bomiense C.Y.Wu & D.D.Tao
- Thesium brachyanthum A.W.Hill
- Thesium brachygyne Schltr.
- Thesium brachyphyllum Boiss.
- Thesium brachystegium Post
- Thesium brachystylum A.W.Hill
- Thesium brasiliense A.DC.
- Thesium brevibarbatum Pilg.
- Thesium brevibracteatum Sumnev.
- Thesium breyeri N.E.Br.
- Thesium bundiense Hilliard
- Thesium burchellii A.W.Hill
- Thesium burkei A.W.Hill
- Thesium canariense (Stearn) J.C.Manning & F.Forest
- Thesium capitatum L.
- Thesium capitellatum A.DC.
- Thesium capituliflorum Sond.
- Thesium carinatum A.DC.
- Thesium catalaunicum Pedrol & M.Laínz
- Thesium cathaicum Hendrych
- Thesium celatum N.E.Br.
- Thesium chimanimaniense Brenan
- Thesium chinense Turcz.
- Thesium cilicicum Hausskn. ex Bornm.
- Thesium cinereum A.W.Hill
- Thesium commutatum Sond.
- Thesium compressum Boiss. & Heldr.
- Thesium confine Sond.
- Thesium congestum R.A.Dyer
- Thesium coriarium A.W.Hill
- Thesium cornigerum A.W.Hill
- Thesium corsalpinum Hendrych
- Thesium costatum A.W.Hill
- Thesium corymbuligerum Sond.
- Thesium crassifolium Sond.
- Thesium crassipes Robyns & Lawalrée
- Thesium cruciatum A.W.Hill
- Thesium cupressoides A.W.Hill
- Thesium cuspidatum A.W.Hill
- Thesium cymosum A.W.Hill
- Thesium cystoseiroides Baker
- Thesium cytisoides A.W.Hill
- Thesium davidsoniae Brenan
- Thesium decaryanum Cavaco & Keraudren
- Thesium deceptum N.E.Br.
- Thesium decipiens Hilliard & B.L.Burtt
- Thesium densiflorum A.DC.
- Thesium densum N.E.Br.
- Thesium dinteri A.W.Hill
- Thesium disciflorum A.W.Hill
- Thesium disparile N.E.Br.
- Thesium dissitiflorum Schltr.
- Thesium dissitum N.E.Br.
- Thesium divaricatum Jan ex Mert. & Koch, raskrečeni lanak
- Thesium diversifolium Sond.
- Thesium dmmagiae Zhigila, Verboom & Muasya
- Thesium dolichomeres Brenan
- Thesium dollineri Murb. ex Velen., Dollinerov lanak
- Thesium doloense Pilg.
- Thesium dumale N.E.Br.
- Thesium durum Hilliard & B.L.Burtt
- Thesium ebracteatum Hayne
- Thesium ecklonianum Sond.
- Thesium elatius Sond.
- Thesium emodi Hendrych
- Thesium equisetoides Welw. ex Hiern
- Thesium ericaefolium A.DC.
- Thesium erythronicum Pamp.
- Thesium euphorbioides P.J.Bergius
- Thesium euphrasioides A.DC.
- Thesium exile N.E.Br.
- Thesium fallax Schltr.
- Thesium fanshawei Hilliard
- Thesium fastigiatum A.W.Hill
- Thesium ferganense Bobrov
- Thesium filipes A.W.Hill
- Thesium fimbriatum A.W.Hill
- Thesium flexuosum A.DC.
- Thesium foliosum A.DC.
- Thesium fragile L.f.
- Thesium frisea L.
- Thesium fruticosum A.W.Hill
- Thesium fruticulosum (A.W.Hill) J.C.Manning & F.Forest
- Thesium fulvum A.W.Hill
- Thesium funale L.
- Thesium fuscum A.W.Hill
- Thesium galioides A.DC.
- Thesium germainii Robyns & Lawalrée
- Thesium glaucescens A.W.Hill
- Thesium glomeratum A.W.Hill
- Thesium glomeruliflorum Sond.
- Thesium gnidiaceum A.DC.
- Thesium goetzeanum Engl.
- Thesium gontscharovii Bobrov
- Thesium gracilarioides A.W.Hill
- Thesium gracile A.W.Hill
- Thesium gracilentum N.E.Br.
- Thesium griseum Sond.
- Thesium gypsophiloides A.W.Hill
- Thesium hararense A.G.Mill.
- Thesium helichrysoides A.W.Hill
- Thesium helodes Hilliard
- Thesium hillianum Compton
- Thesium himalense Royle
- Thesium hirsutum A.W.Hill
- Thesium hispidifructum N.Lombard & M.M.Le Roux
- Thesium hispidulum Lam.
- Thesium hockii Robyns & Lawalrée
- Thesium hollandii Compton
- Thesium hookeri Hendrych
- Thesium horridum Pilg.
- Thesium humbertii Cavaco & Keraudren
- Thesium humifusum DC.
- Thesium humile Vahl, niski lanak
- Thesium hystricoides A.W.Hill
- Thesium hystrix A.W.Hill
- Thesium imbricatum Thunb.
- Thesium impeditum A.W.Hill
- Thesium impressum Steud. ex A.DC.
- Thesium indicum Hendrych
- Thesium infundibulare N.Visser & M.M.le Roux
- Thesium inhambanense Hilliard
- Thesium inonoense Hilliard
- Thesium inversum N.E.Br.
- Thesium italicum A.DC.
- Thesium jarmilae Hendrych
- Thesium jeaniae Brenan
- Thesium junceum Bernh. ex Krauss
- Thesium juncifolium A.DC.
- Thesium junodii A.W.Hill
- Thesium karooicum Compton
- Thesium katangense Robyns & Lawalrée
- Thesium kernerianum Simonk.
- Thesium kilimandscharicum Engl.
- Thesium kotschyanum Boiss.
- Thesium krymense Romo, Didukh & Borat.
- Thesium kyrnosum Hendrych
- Thesium lacinulatum A.W.Hill
- Thesium laetum Robyns & Lawalrée
- Thesium leandrianum Cavaco & Keraudren
- Thesium leptocaule Sond.
- Thesium leptostachyum A.DC.
- Thesium lesliei N.E.Br.
- Thesium leucanthum Gilg
- Thesium lewallei Lawalrée
- Thesium libanoticum Ehrenb. ex A.DC.
- Thesium libericum Hepper & Keay
- Thesium linophyllon L., obični lanak
- Thesium lisae-mariae Stauffer ex Markgr.
- Thesium lisowskii Lawalrée
- Thesium litoreum Brenan
- Thesium lobelioides A.DC.
- Thesium longicaule Zhigila, Verboom & Muasya
- Thesium longiflorum Hand.-Mazz.
- Thesium longifolium Turcz.
- Thesium longiperianthium H.H.Xu & W.Jun
- Thesium lopollense Hiern
- Thesium luembense Robyns & Lawalrée
- Thesium lycopodioides Gilg
- Thesium lynesii Robyns & Lawalrée
- Thesium macedonicum Hendrych
- Thesium macranthum Fenzl
- Thesium macrostachyum A.DC.
- Thesium madagascariense A.DC.
- Thesium magalismontanum Sond.
- Thesium magnifructum Hilliard
- Thesium malaissei Lawalrée
- Thesium manikense Robyns & Lawalrée
- Thesium maritimum C.A.Mey.
- Thesium masukense A.W.Hill
- Thesium matteii Chiov.
- Thesium mauritanicum Batt.
- Thesium megalocarpum A.W.Hill
- Thesium microcarpum A.DC.
- Thesium microcephalum A.W.Hill
- Thesium micromeria A.DC.
- Thesium microphyllum Robyns & Lawalrée
- Thesium micropogon A.DC.
- Thesium minkwitzianum B.Fedtsch.
- Thesium minus (A.W.Hill) J.C.Manning & F.Forest
- Thesium mossii N.E.Br.
- Thesium mukense A.W.Hill
- Thesium multicaule Ledeb.
- Thesium multiramulosum Pilg.
- Thesium myriocladum Baker ex A.W.Hill
- Thesium namaquense Schltr.
- Thesium natalense Sond.
- Thesium nationiae A.W.Hill
- Thesium nautimontanum M.A.García, Nickrent & Mucina
- Thesium neoprostratum Zhigila, Verboom & Muasya
- Thesium nigricans Rendle
- Thesium nigromontanum Sond.
- Thesium nigroperianthum Zhigila, Verboom & Muasya
- Thesium nigrum A.W.Hill
- Thesium nudicaule A.W.Hill
- Thesium nutans Robyns & Lawalrée
- Thesium occidentale A.W.Hill
- Thesium oreogetum Hendrych
- Thesium oresigenum Compton
- Thesium orgadophilum P.C.Tam
- Thesium orientale A.W.Hill
- Thesium ovatifolium N.Lombard & M.M.le Roux
- Thesium pallidum A.DC.
- Thesium palmense P.Pérez & P.Sosa
- Thesium panganense Polhill
- Thesium paniculatum L.
- Thesium parnassi A.DC., Parnasov lanak
- Thesium paronychioides Sond.
- Thesium passerinoides Robyns & Lawalrée
- Thesium patersonae A.W.Hill
- Thesium patulum A.W.Hill
- Thesium pawlowskianum Lawalrée
- Thesium penicillatum A.W.Hill
- Thesium perrieri Cavaco & Keraudren
- Thesium phyllostachyum Sond.
- Thesium pilosum A.W.Hill
- Thesium pinifolium A.DC.
- Thesium pleuroloma A.W.Hill
- Thesium polycephalum Schltr.
- Thesium polygaloides A.W.Hill
- Thesium procerum N.E.Br.
- Thesium procumbens C.A.Mey.
- Thesium prostratum A.W.Hill
- Thesium pseudocystoseiroides Cavaco & Keraudren
- Thesium pseudovirgatum Levyns
- Thesium psilotocladum Svent.
- Thesium psilotoides Hance
- Thesium pubescens A.DC.
- Thesium pungens A.W.Hill
- Thesium pycnanthum Schltr.
- Thesium pygmaeum Hilliard
- Thesium pyrenaicum Pourr., pirenejski lanak
- Thesium quarrei Robyns & Lawalrée
- Thesium quartzicolum Zhigila, Verboom & Muasya
- Thesium quinqueflorum Sond.
- Thesium racemosum Bernh. ex Krauss
- Thesium radicans Hochst.
- Thesium ramosissimum Bobrov
- Thesium ramosoides Hendrych
- Thesium ramosum Hayne
- Thesium rariflorum Sond.
- Thesium rasum (A.W.Hill) N.E.Br.
- Thesium rectangulum Welw. ex Hiern
- Thesium reekmansii Lawalrée
- Thesium refractum C.A.Mey.
- Thesium remotebracteatum C.Y.Wu & D.D.Tao
- Thesium repandum A.W.Hill
- Thesium repens Ledeb.
- Thesium resedoides A.W.Hill
- Thesium resinifolium N.E.Br.
- Thesium retamoides (A.Santos) J.C.Manning & F.Forest
- Thesium rhizomatum Zhigila, Verboom & Muasya
- Thesium rigidum Sond.
- Thesium robynsii Lawalrée
- Thesium rogersii A.W.Hill
- Thesium rostratum Mert. & Koch, kljunav lanak
- Thesium rufescens A.W.Hill
- Thesium rupestre Ledeb.
- Thesium sawae Zhigila, Verboom & Muasya
- Thesium saxatile Turcz.
- Thesium scabriflorum P.H.Davis
- Thesium scabrum L.
- Thesium scandens E.Mey. ex A.DC.
- Thesium schaijesii Lawalrée
- Thesium schliebenii Pilg.
- Thesium schmitzii Robyns & Lawalrée
- Thesium schumannianum Schltr.
- Thesium schweinfurthii Engl.
- Thesium scirpoides A.W.Hill
- Thesium selagineum A.DC.
- Thesium semotum N.E.Br.
- Thesium sertulariastrum A.W.Hill
- Thesium setulosum Robyns & Lawalrée
- Thesium singulare Hilliard
- Thesium sommieri Hendrych
- Thesium sonderianum Schltr.
- Thesium spicatum L.
- Thesium spinosum L.f.
- Thesium spinulosum A.DC.
- Thesium squarrosum L.f.
- Thesium stellerioides Jaub. & Spach
- Thesium stirtonii Zhigila, Verboom & Muasya
- Thesium strictum P.J.Bergius
- Thesium strigulosum A.DC.
- Thesium strigulosum Welw. ex Hiern
- Thesium stuhlmannii Engl.
- Thesium subaphyllum Engl.
- Thesium subnudum Sond.
- Thesium subsimile N.E.Br.
- Thesium subsucculentum (Kämmer) J.C.Manning & F.Forest
- Thesium susannae A.W.Hill
- Thesium symoensii Lawalrée
- Thesium szowitsii A.DC.
- Thesium tauricolum Boiss. & Hausskn.
- Thesium tenuissimum Hook.f.
- Thesium tepuiense Steyerm.
- Thesium tetragonum A.W.Hill
- Thesium thamnus Robyns & Lawalrée
- Thesium thomsonii Hendrych
- Thesium tongolicum Hendrych
- Thesium translucens A.W.Hill
- Thesium transvaalense Schltr.
- Thesium triflorum Thunb. ex L.f.
- Thesium triste A.W.Hill
- Thesium tuvense Krasnob.
- Thesium umbelliferum A.W.Hill
- Thesium unicaule Haines
- Thesium unyikense Engl.
- Thesium urceolatum A.W.Hill
- Thesium ussanguense Engl.
- Thesium utile A.W.Hill
- Thesium vahrmeijeri Brenan
- Thesium vimineum Robyns & Lawalrée
- Thesium virens E.Mey. ex A.DC.
- Thesium virgatum Lam.
- Thesium viride A.W.Hill
- Thesium viridifolium Levyns
- Thesium vlachorum Aldén
- Thesium welwitschii Hiern
- Thesium whitehillense Compton
- Thesium whyteanum Rendle
- Thesium wightianum Wall.
- Thesium wilczekianum Lawalrée
- Thesium xerophyticum A.W.Hill
- Thesium zeyheri A.DC.
- Thesium ×hybridum Beck
- Thesium ×vandasii Rohlena