Paramacrauchenia

Paramacrauchenia; Temporal range: Early Miocene (Colhuehuapian-Santacrucian); ~21–17.8 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
	Left dentary fragment of Paramacrauchenia scamnata
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	†Litopterna
Family:	†Proterotheriidae
Subfamily:	†Proterotheriinae
Genus:	†Paramacrauchenia; Bordas, 1939
Type species
	†Paramacrauchenia scamnata; (Ameghino, 1902)
Other species
	†P. inexspectata Soria, 2001

Paramacrauchenia is an extinct genus of proterotheriid litopterns from the Early Miocene of what is now Argentina and Chile. Its fossils have been found in the Sarmiento and Santa Cruz Formations of Argentina and Chile.^[1]^[2]^[3]

Taxonomy

Paramacrauchenia was originally assigned to the family Macraucheniidae,^[4] however, recent studies now consider it a member of the family Proterotheriidae, where it is found to be a derived member, closely related to Lambdaconus.^[5]^[6]

Below is a phylogenetic tree of the Proterotheriidae, based on the work of McGrath et al. 2020.^[5]

Proterotheriidae

Megadolodinae

	Megadolodus

	Bounodus

Indaleciidae

	Indalecia

	Adiantoides

Paramacrauchenia

Anisolambda

	Lambdaconus suinus

	Lambdaconus inaequifacies

Lambdaconus lacerum

Paranisolambda

Picturotherium

	Neodolodus

	Protheosodon

Anisolophus floweri

Anisolophus australis

	Anisolophus minusculus

	Prolicaphrium

References

^ Kramarz, Alejandro G.; Bond, Mariano (2005). "Los Litopterna (Mammalia) de la Formación Pinturas, Mioceno Temprano-Medio de Patagonia". Ameghiniana (in Spanish). 42 (3): 611–625. ISSN 1851-8044.
^ Bostelmann, J. Enrique; Roux, Jacobous P. Le; Vásquez, Ana; Gutiérrez, Néstor M.; Oyarzún, José Luis; Carreño, Catalina; Torres, Teresa; Otero, Rodrigo; Llanos, Andrea; Fanning, C. Mark; Hervé, Francisco (2013-07-25). "Burdigalian deposits of the Santa Cruz Formation in the Sierra Baguales, Austral (Magallanes) Basin: Age, depositional environment and vertebrate fossils". Andean Geology. 40 (3): 458–489. doi:10.5027/andgeoV40n3-a04. hdl:10533/138811. ISSN 0718-7106.
^ Bostelmann, Enrique; Le Roux, Jacobus P.; Vasquez, Ana; Gutiérrez, Nestor; Oyarzún, José Luis; Carreño, Catalina; Torres, Teresa; Otero, Rodrigo; Llanos, Andrea; Fanning, C. Mark; Nielsen, Sven. N.; Hervé, Francisco (2012). A revised lithostratigraphy of the Sierra Baguales, Magallanes Basin (PDF). XIII Congeso Geológico Chileno. pp. 698–700. Archived from the original (PDF) on November 3, 2018.
^ Carroll, Robert L. (Robert Lynn) 1938- (1988). Vertebrate paleontology and evolution (PDF). New York, N.Y.: W.H. Freeman and Company. ISBN 0-521-47232-6. OCLC 14967288.{{cite book}}: CS1 maint: numeric names: authors list (link)
^ ^a ^b Andrew J. McGrath; Federico Anaya; Darin A. Croft (2020). "New proterotheriids (Litopterna, Mammalia) from the middle Miocene of Quebrada Honda, Bolivia, and trends in diversity and body size of proterotheriid and macraucheniid litopterns". Ameghiniana. 57 (2): 159–188. doi:10.5710/AMGH.03.03.2020.3268. S2CID 216236954.
^ Schmidt, Gabriela I. (2013-04-25). Litopterna y Notoungulata (Mammalia) de la formación Ituzaingó (Mioceno tardío-Plioceno) de la provincia de Entre Ríos: sistemática, bioestratigrafía y paleobiogeografía (Tesis thesis) (in Spanish). Universidad Nacional de La Plata.

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[Picturotherium-1] Kramarz, Alejandro G.; Bond, Mariano (2005). "Los Litopterna (Mammalia) de la Formación Pinturas, Mioceno Temprano-Medio de Patagonia". Ameghiniana (in Spanish). 42 (3): 611–625. ISSN 1851-8044.

[Bostelmannetal2013-2] Bostelmann, J. Enrique; Roux, Jacobous P. Le; Vásquez, Ana; Gutiérrez, Néstor M.; Oyarzún, José Luis; Carreño, Catalina; Torres, Teresa; Otero, Rodrigo; Llanos, Andrea; Fanning, C. Mark; Hervé, Francisco (2013-07-25). "Burdigalian deposits of the Santa Cruz Formation in the Sierra Baguales, Austral (Magallanes) Basin: Age, depositional environment and vertebrate fossils". Andean Geology. 40 (3): 458–489. doi:10.5027/andgeoV40n3-a04. hdl:10533/138811. ISSN 0718-7106.

[Bost2012-3] Bostelmann, Enrique; Le Roux, Jacobus P.; Vasquez, Ana; Gutiérrez, Nestor; Oyarzún, José Luis; Carreño, Catalina; Torres, Teresa; Otero, Rodrigo; Llanos, Andrea; Fanning, C. Mark; Nielsen, Sven. N.; Hervé, Francisco (2012). A revised lithostratigraphy of the Sierra Baguales, Magallanes Basin (PDF). XIII Congeso Geológico Chileno. pp. 698–700. Archived from the original (PDF) on November 3, 2018.

[4] Carroll, Robert L. (Robert Lynn) 1938- (1988). Vertebrate paleontology and evolution (PDF). New York, N.Y.: W.H. Freeman and Company. ISBN 0-521-47232-6. OCLC 14967288.{{cite book}}: CS1 maint: numeric names: authors list (link)

[mcgrath2020-5] Andrew J. McGrath; Federico Anaya; Darin A. Croft (2020). "New proterotheriids (Litopterna, Mammalia) from the middle Miocene of Quebrada Honda, Bolivia, and trends in diversity and body size of proterotheriid and macraucheniid litopterns". Ameghiniana. 57 (2): 159–188. doi:10.5710/AMGH.03.03.2020.3268. S2CID 216236954.

[6] Schmidt, Gabriela I. (2013-04-25). Litopterna y Notoungulata (Mammalia) de la formación Ituzaingó (Mioceno tardío-Plioceno) de la provincia de Entre Ríos: sistemática, bioestratigrafía y paleobiogeografía (Tesis thesis) (in Spanish). Universidad Nacional de La Plata.

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