Actitis

Actitis
	Common sandpiper, Actitis hypoleucos
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Order:	Charadriiformes
Family:	Scolopacidae
Genus:	Actitis; Illiger, 1811
Type species
	Tringa hypoleucos (common sandpiper); Linnaeus, 1758
Species
	Actitis hypoleucos; Actitis macularius

Actitis is a small genus of waders, comprising just two very similar bird species.

Taxonomy

The genus Actitis was introduced in 1811 by the German zoologist Johann Illiger.^[1] The genus name is from Ancient Greek aktites, "coast-dweller" from akte, "coast".^[2] The type species is the common sandpiper.^[3] The genus is sister to the genus Tringa that contains the shanks and the tattlers.^[4]

The genus contains two species:^[5]

Genus **Actitis** – Illiger, 1811 – two species
Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Common sandpiper	Actitis hypoleucos (Linnaeus, 1758)	Eurasia	Size: Habitat: Diet:	LC
Spotted sandpiper	Actitis macularius (Linnaeus, 1766)	North America	Size: Habitat: Diet:	LC

Description

These are both small migratory waders, greyish brown on top and white underneath, with a distinctive stiff-winged flight low over the water. The plumages are very similar, apart from spotted sandpipers' distinctive breeding plumage, and suspected out-of-range vagrants must be carefully observed for identification to species.

Both species have short yellow or yellowish legs and a medium bill. These are not gregarious birds and are seldom seen in large flocks.

They nest on the ground, and their habitat is near fresh water. These birds forage on the ground or in water, picking up food by sight. They may also catch insects in flight. They eat insects, crustaceans and other invertebrates.

Actitis is part of the shank-tattler-phalarope clade and less closely related to the calidrid sandpipers.^[6] Based on the degree of DNA sequence divergence and putative shank and phalarope fossils from around the Oligocene/Miocene boundary some 23–22 million years ago, presumably Actitis diverged from its closest relatives in the Late Oligocene; given the much higher diversity of the prehistoric members of the group in Eurasia it is likely that they originated there, possibly being isolated as the remains of the Turgai Sea dried up, which happened just around this time.^[7]

The Late Pliocene fossil described as Actitis balcanica ^[8] appears to be actually from some indeterminate charadriid.^[9]

Footnotes

^ Illiger, Johann Karl Wilhelm (1811). Prodromus systematis mammalium et avium (in Latin). Berolini [Berlin]: Sumptibus C. Salfeld. p. 262.
^ Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. p. 31. ISBN 978-1-4081-2501-4.
^ Peters, James Lee, ed. (1934). Check-List of Birds of the World. Vol. 2. Cambridge, Massachusetts: Harvard University Press. p. 269.
^ Gibson, Rosemary; Baker, Allan (2012). "Multiple gene sequences resolve phylogenetic relationships in the shorebird suborder Scolopaci (Aves: Charadriiformes)". Molecular Phylogenetics and Evolution. 64 (1): 66–72. Bibcode:2012MolPE..64...66G. doi:10.1016/j.ympev.2012.03.008. PMID 22491071.
^ Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (July 2021). "Sandpipers, snipes, coursers". IOC World Bird List Version 11.2. International Ornithologists' Union. Retrieved 22 November 2021.
^ Paton et al. (2003), Thomas et al. (2004)
^ Mlíkovský (2002), Paton et al. (2003)
^ Boev, Z. 1998. Actitis balcanica sp. n. - a Late Pliocene Sandpiper (Aves: Scolopacidae) from Bulgaria. - Historia naturalis bulgarica, 9: 71–77.
^ Mlíkovský (2002)

References

Mlíkovský, Jirí (2002): Cenozoic Birds of the World, Part 1: Europe. Ninox Press, Prague. ISBN 80-901105-3-8
Paton, Tara A.; Baker, Allan J.; Groth, J. G.; Barrowclough, G. F. (2003). "RAG-1 sequences resolve phylogenetic relationships within charadriiform birds". Molecular Phylogenetics and Evolution. 29 (2): 268–278. Bibcode:2003MolPE..29..268P. doi:10.1016/S1055-7903(03)00098-8. PMID 13678682.
Thomas, Gavin H.; Wills, Matthew A.; Székely, Tamás (2004). "A supertree approach to shorebird phylogeny". BMC Evolutionary Biology. 4: 28. doi:10.1186/1471-2148-4-28. PMC 515296. PMID 15329156. Supplementary Material

[1] Illiger, Johann Karl Wilhelm (1811). Prodromus systematis mammalium et avium (in Latin). Berolini [Berlin]: Sumptibus C. Salfeld. p. 262.

[job-2] Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. p. 31. ISBN 978-1-4081-2501-4.

[3] Peters, James Lee, ed. (1934). Check-List of Birds of the World. Vol. 2. Cambridge, Massachusetts: Harvard University Press. p. 269.

[4] Gibson, Rosemary; Baker, Allan (2012). "Multiple gene sequences resolve phylogenetic relationships in the shorebird suborder Scolopaci (Aves: Charadriiformes)". Molecular Phylogenetics and Evolution. 64 (1): 66–72. Bibcode:2012MolPE..64...66G. doi:10.1016/j.ympev.2012.03.008. PMID 22491071.

[ioc-5] Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (July 2021). "Sandpipers, snipes, coursers". IOC World Bird List Version 11.2. International Ornithologists' Union. Retrieved 22 November 2021.

[6] Paton et al. (2003), Thomas et al. (2004)

[7] Mlíkovský (2002), Paton et al. (2003)

[8] Boev, Z. 1998. Actitis balcanica sp. n. - a Late Pliocene Sandpiper (Aves: Scolopacidae) from Bulgaria. - Historia naturalis bulgarica, 9: 71–77.

[9] Mlíkovský (2002)

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