Sri Lanka is a tropical island situated close to the southern tip of India. It is situated in the middle of Indian Ocean. Because of being an island, Sri Lanka has rich endemic terrestrial and freshwater fauna, including vertebrates and several invertebrates.
Amphibian
editAmphibians are ectothermic vertebrates of the class Amphibia. They have soft glandular skin and live in all habitats of the world except for the ice caps. They complete an amphibious lifestyle where larval stages live in water and adults live on or closer to land. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators.
Sri Lanka is host to over 120 species of amphibians, of which over 90 species are endemic to the country.[1] The 85% of endemicity ratio makes Sri Lanka the country which has the highest amphibian endemism in Asia.[2] During the past decade many more new amphibians have been found in Sri Lanka.[3]
The first amphibian review in Sri Lanka in 1957 identified 35 species.[4] In 1996 the number of amphibian species rose to 53 based on research of museum collections and also a field survey. More than 250 species were proposed based on this field survey by Rohan Pethiyagoda and Kelum Manamendra-Arachchi in 1998. However Madhava Meegaskumbura et al. revised the number to around 140 species,[5] and the discovery of over "new" 100 species has been criticised.[6] As at 2019, 122 descriptions of amphibian species have been published, with 113 endemics.[7][8] Three caecilian species has been identified with one undescribed species.
Sri Lanka harbours three endemic genera, Adenomus, Nannophrys, and Lankanectes.[3] Most of the new species are of the genus Philautus which was assigned to genus Pseudophilautus recently. Hence, there are no amphibians of the genus Philautus (sensu stricto) in Sri Lanka. Pseudophilautus pardus and P. maia, the species known only from collections made prior to 1876 are described as new species in 2007, but both are extinct.[9] In April 2015, Mendis Wickrremasinghe et al. described another endemic Pseudophilautus species P. dilmah. In January 2019, the new species, P. conniffae was discovered in southern Sri Lanka.[10]
Sri Lanka has the highest percentage of extinct and threatened amphibian species in Asia.[2] In the 20th century the country has lost 20% of its amphibians and more than half of the remaining species are on the verge of extinction. Of the world's 34 amphibian fauna that have gone extinct in the last 500 years, 19 are from Sri Lanka.[3] Habitat loss is attributed as the main cause of threats while fragmentation, use of pesticides, and air pollution are among others.
Amphibian diversity of Sri Lanka
editLow vulnerability | Threatened | Extinct | Insufficient data | |||||||||||||||||||||
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Order Anura: frogs
editCommon name | Species
(authority) |
Endemicity | Status |
---|---|---|---|
Family Bufonidae: true toads | |||
Kandyan dwarf toad | Adenomus kandianus (Günther, 1872) |
Endemic | [11] |
Kelaart's dwarf toad | Adenomus kelaartii (Günther, 1858) |
Endemic | [12] |
Kotagama's dwarf toad | Duttaphrynus kotagamai Fernando & Dayawansa, 1994 |
Endemic | [13] |
Common Indian toad | Duttaphrynus melanostictus Schneider, 1799 |
[14] | |
Noellert's toad | Duttaphrynus noellerti Manamendra-Arachchi & Pethiyagoda, 1998 |
Endemic | [15] |
Ferguson's toad | Duttaphrynus scaber Schneider, 1799 |
[16] | |
Marbled toad | Duttaphrynus stomaticus Lütken, 1862 |
[16] | |
Family Dicroglossidae: fork-tongued frogs | |||
Indian skipper frog | Euphlyctis mudigere (Joshy, Alam, Kurabayashi, Sumida, and Kuramoto, 2009) |
[17] | |
Green pond frog | Euphlyctis hexadactylus (Lesson, 1834) |
[18] | |
Jerdon's bullfrog | Hoplobatrachus crassus (Jerdon, 1853) |
[19] | |
Sri Lanka rock frog | Nannophrys ceylonensis (Günther, 1868) |
Endemic | [20] |
Gunther's streamlined frog | Nannophrys guentheri Boulenger, 1882 |
Endemic | [21] |
Kirtisinghe's rock frog | Nannophrys marmorata Kirtisinghe, 1946 |
Endemic | [22] |
Nayak rock frog | Nannophrys naeyakai Fernando, Wickramasingha & Rodrigo, 2007 |
Endemic | [23] |
Indian burrowing frog | Sphaerotheca breviceps (Schneider, 1799) |
[24] | |
Sri Lanka burrowing frog | Sphaerotheca rolandae (Dubois, 1983) |
Endemic | [25] |
Common Indian cricket frog | Minervarya agricola (Jerdon, 1853) |
[26] | |
Montane frog | Minervarya greenii (Boulenger, 1904) |
Endemic | [27] |
Kirtisinghe's frog | Minervarya kirtisinghei Manamendra-Arachchi & Gabadage, 1994 |
Endemic | [28] |
Family Microhylidae: narrow-mouthed frogs | |||
Karunaratne's narrow-mouthed frog | Microhyla karunaratnei Fernando & Siriwardhane, 1996 |
Endemic | [29] |
Mihintale red narrow-mouthed frog | Microhyla mihintalei Wijayathilaka, Garg, Senevirathne, Karunarathna, Biju & Meegaskumbura, 2016 |
Endemic | |
Ornate narrow-mouthed frog | Microhyla ornata (Duméril & Bibron, 1841) |
[30] | |
Sri Lanka rice frog | Microhyla zeylanica Parker & Hill, 1949 |
Endemic | [31] |
Sri Lankan painted frog | Uperodon taprobanicus (Parker, 1934) |
[32] | |
Nagao's pug-snouted frog | Uperodon nagaoi (Manamendra-Arachchi & Pethiyagoda, 2001) |
Endemic | [33] |
Sri Lanka dot frog | Uperodon obscurus (Günther, 1864) |
Endemic | [34] |
Half-webbed pug-snouted frog | Uperodon palmatus (Parker, 1934) |
Endemic | [35] |
Rohan's globular frog | Uperodon rohani
(Garg S, Senevirathne G, Wijayathilaka N, Phuge S, Deuti K, Manamendra-Arachchi K, Meegaskumbura M, Biju SD. 2018) |
Endemic | [36] |
Marbled balloon frog | Uperodon systoma (Schneider, 1799) |
[37] | |
Family Ranidae: true frogs | |||
Sri Lankan golden-backed frog | Indosylvirana serendipi Biju et al., 2014 |
Endemic | [38] |
Gravenhorst's frog | Hydrophylax gracilis Gravenhorst, 1829 |
Endemic | [39] |
Bronzed frog | Indosylvirana temporalis (Günther, 1864) |
Endemic | [40] |
Family Nyctibatrachidae: robust frogs | |||
Corrugated water frog | Lankanectes corrugatus (Peters, 1863) |
Endemic | [41] |
Lankanectes pera Senevirathne et al., 2018 |
Endemic | ||
Family Rhacophoridae: shrub frogs | |||
Pseudophilautus abundus Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [42] | |
Pseudophilautus adspersus (Günther, 1872) |
Endemic | [43] | |
Pseudophilautus alto Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [44] | |
Pseudophilautus asankai Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [45] | |
Pseudophilautus auratus Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [46] | |
Pseudophilautus bambaradeniyai Wickramasinghe et al., 2013 |
Endemic | [47] | |
Pseudophilautus caeruleus Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [48] | |
Pseudophilautus cavirostris (Günther, 1869) |
Endemic | [49] | |
Pseudophilautus conniffae Batuwita, De Silva & Udugampala, 2019 |
Endemic | ||
Pseudophilautus cuspis Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [50] | |
Pseudophilautus dayawansai Wickramasinghe et al. 2013 |
Endemic | [47] | |
Pseudophilautus decoris Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [51] | |
Pseudophilautus dilmah (Wickramasinghe, Bandara, Vidanapathirana, Tennakoon, Samarakoon & Wickramasinghe, 2015) |
Endemic | [52] | |
Pseudophilautus dimbullae (Shreve, 1940) |
Endemic | [52] | |
Pseudophilautus eximius (Shreve, 1940) |
Endemic | [53] | |
Pseudophilautus extirpo Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [54] | |
Pseudophilautus femoralis (Günther, 1864) |
Endemic | [55] | |
Pseudophilautus fergusonianus (Ahl, 1927) |
Endemic | [56] | |
Pseudophilautus folicola Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [57] | |
Pseudophilautus frankenbergi Meegaskumbura & Manamendra-Arachchi, 2005 |
Endemic | [58] | |
Pseudophilautus fulvus Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [59] | |
Pseudophilautus hallidayi Meegaskumbura & Manamendra-Arachchi, 2005 |
Endemic | [60] | |
Pseudophilautus halyi (Boulenger, 1904) |
Endemic | [61] | |
Pseudophilautus hankeni Meegaskumbura & Manamendra-Archchi, 2011 |
Endemic | [60] | |
Pseudophilautus hoipolloi Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [62] | |
Pseudophilautus hypomelas (Günther, 1876) |
Endemic | [63] | |
Pseudophilautus jagathgunawardanai Wickramasinghe et al. 2013 |
Endemic | [47] | |
Pseudophilautus karunarathnai Wickramasinghe et al. 2013 |
Endemic | [47] | |
Pseudophilautus leucorhinus (Lichtenstein, Weinland & Von Martens, 1856) |
Endemic | [64] | |
Pseudophilautus limbus Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [65] | |
Pseudophilautus lunatus Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [66] | |
Pseudophilautus macropus (Günther, 1869) |
Endemic | [67] | |
Pseudophilautus maia (Meegaskumbara et al., 2007) |
Endemic | [67] | |
Pseudophilautus malcolmsmithi (Ahl, 1927) |
Endemic | [68] | |
Pseudophilautus microtympanum (Günther, 1859) |
Endemic | [69] | |
Pseudophilautus mittermeieri Meegaskumbura & Manamendra-Arachchi, 2005 |
Endemic | [70] | |
Pseudophilautus mooreorum Meegaskumbura & Manamendra-Arachchi, 2005 |
Endemic | [71] | |
Pseudophilautus nanus (Günther, 1869) |
Endemic | [72] | |
Pseudophilautus nasutus (Günther, 1869) |
Endemic | [73] | |
Pseudophilautus nemus Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [74] | |
Pseudophilautus newtonjayawardanei Wickramasinghe et al. 2013 |
Endemic | [47] | |
Pseudophilautus ocularis Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [75] | |
Pseudophilautus oxyrhynchus (Günther, 1872) |
Endemic | [76] | |
Pseudophilautus papillosus Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [77] | |
Pseudophilautus pardus (Meegaskumbura, Manamendra-Arachchi, Schneider and Pethiyagoda, 2007) |
Endemic | [67] | |
Pseudophilautus pleurotaenia (Boulenger, 1904) |
Endemic | [78] | |
Pseudophilautus poppiae Meegaskumbura & Manamendra-Arachchi, 2005 |
Endemic | [79] | |
Pseudophilautus popularis Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [80] | |
Pseudophilautus procax Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [81] | |
Pseudophilautus puranappu Wickramasinghe et al. 2013 |
Endemic | [47] | |
Pseudophilautus regius Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [82] | |
Pseudophilautus reticulatus (Günther, 1864) |
Endemic | [83] | |
Pseudophilautus rugatus (Ahl, 1927) |
Endemic | [84] | |
Pseudophilautus rus Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [85] | |
Pseudophilautus samarakoon Wickramasinghe et al. 2013 |
Endemic | [47] | |
Pseudophilautus sarasinorum (Müller, 1887) |
Endemic | [86] | |
Pseudophilautus schmarda (Kelaart, 1854) |
Endemic | [87] | |
Pseudophilautus schneideri Meegaskumbura & Manamendra-Arachchi, 2011 |
Endemic | [88] | |
Pseudophilautus semiruber (Annandale, 1913) |
Endemic | [88] | |
Pseudophilautus silus Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [89] | |
Pseudophilautus silvaticus Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [90] | |
Pseudophilautus simba Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [91] | |
Pseudophilautus singu Meegaskumbura & Manamendra-Arachchi, & Pethiyagoda 2009 |
Endemic | [88] | |
Pseudophilautus sirilwijesundarai Wickramasinghe et al. 2013 |
Endemic | [88] | |
Pseudophilautus sordidus Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [92] | |
Pseudophilautus stellatus (Kelaart, 1853) |
Endemic | [93] | |
Pseudophilautus steineri Meegaskumbura & Manamendra-Arachchi, 2005 |
Endemic | [94] | |
Pseudophilautus stictomerus (Günther, 1876) |
Endemic | [95] | |
Pseudophilautus stuarti Meegaskumbura & Manamendra-Arachchi, 2005 |
Endemic | [96] | |
Pseudophilautus tanu Meegaskumbura & Manamendra-Arachchi, & Pethiyagoda 2009 |
Endemic | [88] | |
Pseudophilautus temporalis (Günther, 1864) |
Endemic | [97] | |
Pseudophilautus variabilis (Günther, 1859) |
Endemic | [98] | |
Pseudophilautus viridis Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [99] | |
Pseudophilautus zal Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [100] | |
Pseudophilautus zimmeri (Ahl, 1927) |
Endemic | [101] | |
Pseudophilautus zorro Manamendra-Arachchi & Pethiyagoda, 2005 |
Endemic | [102] | |
Common hour-glass tree frog | Polypedates cruciger Blyth, 1852 |
Endemic | [103] |
Indian tree frog | Polypedates maculatus (Gray, 1834) |
[104] | |
Ranwella's spined tree frog | Polypedates ranwellai Wickramasinghe, Munindrasasa & Fernando, 2012 |
Endemic | [103] |
Montane hour-glass tree frog | Taruga eques Günther, 1858 |
Endemic | [105] |
Morningside hour-glass tree frog | Taruga fastigo Manamendra-Arachchi & Pethiyagoda, 2001 |
Endemic | [106] |
Long-snouted tree frog | Taruga longinasus (Ahl, 1931) |
Endemic | [107] |
Order Gymnophiona: caecilians
editCommon name | Species
(authority) |
Endemicity | Status |
---|---|---|---|
Family Ichthyophiidae: Asiatic tailed caecilians | |||
Ceylon caecilian | Ichthyophis glutinosus (Linnaeus, 1758) |
Endemic | [108] |
Pattipola caecilian | Ichthyophis orthoplicatus Taylor, 1965 |
Endemic | [109] |
Lesser yellow-banded caecilian | Ichthyophis pseudangularis Taylor, 1965 |
Endemic | [110] |
References
edit- ^ "Species richness of inland and marine fauna in Sri Lanka" (PDF). The 2007 Red List of Threatened Fauna and Flora of Sri Lanka. Colombo, Sri Lanka: IUCN Sri Lanka and the Ministry of Environment and Natural Resources. 2007. p. 3. ISBN 978-955-8177-63-1.
- ^ a b "The state of amphibians in Sri Lanka". iucn.org. IUCN. Archived from the original on 30 April 2010. Retrieved 29 March 2010.
- ^ a b c Bambaradeniya, Channa N.B., ed. (2006). The fauna of Sri Lanka: status of taxonomy, research and conservation (PDF). IUCN Sri Lanka and Government of Sri Lanka. pp. 125–131. ISBN 978-955-8177-51-8.
- ^ "Diversification of Shrub Frogs (Rhacophoridae, Pseudophilautus) in Sri Lanka – Timing and Geographic Context". Novataxa. 23 November 2018. Retrieved 3 February 2019.
- ^ Meegaskumbura, M.; Bossuyt, F.; Pethiyagoda, R.; Manamendra-Arachchi, K.; Bahir, M.; Milinkovitch, M. C.; Schneider, C. J. (2002). "Sri Lanka: An Amphibian Hot Spot" (PDF). Science. 298 (5592): 379. CiteSeerX 10.1.1.477.4345. doi:10.1126/science.298.5592.379. PMID 12376694.
- ^ Chaitra, M. S.; Vasudevan, Karthikeyan; Shanker, Kartik (10 April 2004). "The biodiversity bandwagon: the splitters have it" (PDF). Current Science. 86 (7): 897.
- ^ "Biodiversity of Sri Lanka: Amphibians of Sri Lanka".
- ^ "A new frog species from rapidly dwindling cloud forest streams of Sri Lanka—Lankanectes pera". Zootaxa 4461(4):519-538. Retrieved 25 August 2018.
- ^ Meegaskumbura, Madhava; Manamendra-Arachchi, Kelum; Schneider, Christopher J.; Pethiyagoda, Rohan (25 January 2007). "New species amongst Sri Lanka's extinct shrub frogs (Amphibia: Rhacophoridae: Philautus)" (PDF). Zootaxa. 1397: 1–15. doi:10.11646/zootaxa.1397.1.1. ISSN 1175-5326.
- ^ Udugampala, Sampath; De Silva, Madura; Batuwita, Sudesh (2019). "Description of a new species of Pseudophilautus (Amphibia: Rhacophoridae) from southern Sri Lanka". Journal of Threatened Taxa. 11: 13120–13131. doi:10.11609/jott.3903.11.1.13120-13131. S2CID 92521317. Retrieved 30 January 2019.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Adenomus kandianus". IUCN Red List of Threatened Species. 2020: e.T80161781A80161555. doi:10.2305/IUCN.UK.2020-3.RLTS.T80161781A80161555.en. Retrieved 20 November 2021.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Adenomus kelaartii". IUCN Red List of Threatened Species. 2020: e.T54459A156577899. doi:10.2305/IUCN.UK.2020-3.RLTS.T54459A156577899.en. Retrieved 20 November 2021.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Duttaphrynus kotagamai". IUCN Red List of Threatened Species. 2020: e.T54683A156578105. doi:10.2305/IUCN.UK.2020-3.RLTS.T54683A156578105.en. Retrieved 20 November 2021.
- ^ Peter Paul van Dijk; Djoko Iskandar; Michael Wai Neng Lau; Gu Huiqing; Geng Baorong; Lue Kuangyang; Chou Wenhao; Yuan Zhigang; Bosco Chan; Sushil Dutta; et al. (2004). "Duttaphrynus melanostictus". IUCN Red List of Threatened Species. 2004. Retrieved 28 March 2010.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Duttaphrynus noellerti". IUCN Red List of Threatened Species. 2020: e.T54716A156578206. doi:10.2305/IUCN.UK.2020-3.RLTS.T54716A156578206.en. Retrieved 20 November 2021.
- ^ a b Sushil Dutta, Kelum Manamendra-Arachchi (2004). "Duttaphrynus scaber". IUCN Red List of Threatened Species. 2004: e.T54753A11199889. doi:10.2305/IUCN.UK.2004.RLTS.T54753A11199889.en. Retrieved 20 November 2021.
- ^ Muhammad Sharif Khan; Theodore Papenfuss; Steven Anderson; Nasrullah Rastegar-Pouyani; Sergius Kuzmin; Sushil Dutta; Kelum Manamendra-Arachchi; Mozafar Sharifi (2008). "Euphlyctis cyanophlyctis". IUCN Red List of Threatened Species. 2008. Retrieved 28 March 2010.
- ^ Sushil Dutta, Kelum Manamendra-Arachchi (2004). "Euphlyctis hexadactylus". IUCN Red List of Threatened Species. 2004: e.T58263A11758707. doi:10.2305/IUCN.UK.2004.RLTS.T58263A11758707.en. Retrieved 20 November 2021.
- ^ Anand Padhye, Kelum Manamendra-Arachchi, Anslem de Silva, Sushil Dutta, Tej Kumar Shrestha, Sabitry Bordoloi (2004). "Hoplobatrachus crassus". IUCN Red List of Threatened Species. 2004: e.T58298A11754703. doi:10.2305/IUCN.UK.2004.RLTS.T58298A11754703.en. Retrieved 20 November 2021.
{{cite journal}}
: CS1 maint: multiple names: authors list (link) - ^ IUCN SSC Amphibian Specialist Group (2020). "Nannophrys ceylonensis". IUCN Red List of Threatened Species. 2020: e.T58389A156580160. doi:10.2305/IUCN.UK.2020-3.RLTS.T58389A156580160.en. Retrieved 20 November 2021.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Nannophrys guentheri". IUCN Red List of Threatened Species. 2020: e.T58390A156580255. doi:10.2305/IUCN.UK.2020-3.RLTS.T58390A156580255.en. Retrieved 20 November 2021.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Nannophrys marmorata". IUCN Red List of Threatened Species. 2020: e.T58391A156580316. doi:10.2305/IUCN.UK.2020-3.RLTS.T58391A156580316.en. Retrieved 20 November 2021.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Nannophrys naeyakai". IUCN Red List of Threatened Species. 2020: e.T135839A156588497. doi:10.2305/IUCN.UK.2020-3.RLTS.T135839A156588497.en. Retrieved 20 November 2021.
- ^ Annemarie Ohler; Muhammad Sharif Khan; Peter Paul van Dijk; Guinevere Wogan; Sushil Dutta; Robert Inger; Tej Kumar Shrestha; Kelum Manamendra-Arachchi; Anslem de Silva (2004). "Sphaerotheca breviceps". IUCN Red List of Threatened Species. 2004. Retrieved 28 March 2010.
- ^ Kelum Manamendra-Arachchi, Sushil Dutta (2004). "Sphaerotheca rolandae". IUCN Red List of Threatened Species. 2004: e.T58759A11838218. doi:10.2305/IUCN.UK.2004.RLTS.T58759A11838218.en. Retrieved 20 November 2021.
- ^ Nelsoon Annandale (1919). "Fejervarya syhadrensis". IUCN Red List of Threatened Species. 1919. Retrieved 28 March 2010.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Minervarya greenii". IUCN Red List of Threatened Species. 2020: e.T58271A156579645. doi:10.2305/IUCN.UK.2020-3.RLTS.T58271A156579645.en. Retrieved 20 November 2021.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Minervarya kirtisinghei". IUCN Red List of Threatened Species. 2020: e.T58274A156579783. doi:10.2305/IUCN.UK.2020-3.RLTS.T58274A156579783.en. Retrieved 20 November 2021.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Microhyla karunaratnei". IUCN Red List of Threatened Species. 2020: e.T57883A156578570. doi:10.2305/IUCN.UK.2020-3.RLTS.T57883A156578570.en. Retrieved 20 November 2021.
- ^ Dutta, S.; Kumar Shrestha, T.; Manamendra-Arachchi, K.; Khan, M.S.; Roy, D. (2008). "Microhyla ornata". IUCN Red List of Threatened Species. 2008: e.T57886A11686884. doi:10.2305/IUCN.UK.2008.RLTS.T57886A11686884.en. Retrieved 20 November 2021.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Microhyla zeylanica". IUCN Red List of Threatened Species. 2020: e.T57895A156578682. doi:10.2305/IUCN.UK.2020-3.RLTS.T57895A156578682.en. Retrieved 20 November 2021.
- ^ Inger, R.F.; Gour-Broome, V.A.; Manamendra-Arachchi, K.; de Silva, A.; Dutta, S. (2016). "Uperodon taprobanicus". IUCN Red List of Threatened Species. 2016: e.T57858A91639191. doi:10.2305/IUCN.UK.2016-1.RLTS.T57858A91639191.en. Retrieved 20 November 2021.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Uperodon nagaoi". IUCN Red List of Threatened Species. 2020: e.T57988A156578796. doi:10.2305/IUCN.UK.2020-3.RLTS.T57988A156578796.en. Retrieved 20 November 2021.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Uperodon obscurus". IUCN Red List of Threatened Species. 2020: e.T57989A156578879. doi:10.2305/IUCN.UK.2020-3.RLTS.T57989A156578879.en. Retrieved 20 November 2021.
- ^ IUCN SSC Amphibian Specialist Group (2020). "Uperodon palmatus". IUCN Red List of Threatened Species. 2020: e.T57990A156578985. doi:10.2305/IUCN.UK.2020-3.RLTS.T57990A156578985.en. Retrieved 20 November 2021.
- ^ Dutta, S.; Manamendra-Arachchi, K. (2016). "Uperodon variegatus". IUCN Red List of Threatened Species. 2016: e.T57992A91636474. doi:10.2305/IUCN.UK.2016-1.RLTS.T57992A91636474.en. Retrieved 20 November 2021.
- ^ Das, I.; Dutta, S.; Manamendra-Arachchi, K.; de Silva, A.; Sharif Khan, M. (2004). "Uperodon systoma". IUCN Red List of Threatened Species. 2004. Retrieved 28 March 2010.
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